Deadline: April 15, 2022. Course commencing 10-16 June 2022.

The course will be held at Tjärnö Marine Laboratory, University of Gothenburg, outside Strömstad, Sweden.

The course is free of charge and lodging is also covered for. Students will have to cover their own travel and costs for food. The course will take place 10–16 June 2022, with arrival date June 9th and departure day June 17th.

Through lectures and laboratory exercises the course will explore the role of variation and extremes in ecology and ecological physiology. Topics addressed will include the pervasive effects of Jensen’s inequality (“the fallacy of the average”), the statistics of extremes and how to calculate and interpret them, and the importance of mechanistic models in the exploration of variation and extremes.

The course is aimed at graduate students and postdoctoral fellows in ecology, ecological physiology, and related areas of biology and environmental science. Students are encouraged to bring to the class their own data and their experiences with the topic. The course corresponds to 4 ECT.


The application should include a short motivation (<1 page) and a brief CV. Submit to the University of Gothenburg course portal.

For more information, please see the full course information below.